Publications

SEAMARCO distributes knowledge on marine animals in the following ways:

Studies

Research conducted by SEAMARCO is published in internationally renowned scientific journals, so that the results can be used world-wide for the protection and management of marine animals. The following studies have been published:

Showing 16 categories
YearTitleAuthorsPublished inCategory
1983"Behavioral and physiological responses of captive beluga whales to playbacks of drilling noise"Thomas, J.A., Kastelein, R.A.J. Acoust. Soc. Am. 74(SI),S 54 (A)Acoustics / Behavioral response studies (Effect of sound on behavior)
1990"Behavior and blood catecholamines of captive Belugas during playbacks of noise from an oil drilling platform"Thomas, J.A., Kastelein, R.A., Awbrey, F.T.Zoo Biology 9,393-402Acoustics / Behavioral response studies (Effect of sound on behavior)
1995"The effects of acoustic alarms on Harbour porpoise (Phocoena phocoena) behaviour"Kastelein, R.A., Goodson, A.D., Lien, J., de Haan, D.Harbour porpoises, laboratory studies to reduce bycatch (Eds Nachtigall, P.E., Lien, J., Au, W.W.L. and Read, A.J.). De Spil Publishers, Woerden, The Netherlands, 157-167Acoustics / Behavioral response studies (Effect of sound on behavior)
1997"The response of a harbour porpoise (Phocoena phocoena) to nets of various mesh sizes, with and without deterring sound"Kastelein, R.A., de Haan, D., Staal, C., Goodson, A.D.The biology of the harbour porpoise (Eds. Read, A.J., Wiepkema, P.R. and Nachtigall, P.E.). De Spil Publishers, Woerden, The Netherlands, 385-409Acoustics / Behavioral response studies (Effect of sound on behavior)
1997"The effects of various sounds on a harbour porpoise (Phocoena phocoena)"Kastelein, R.A., de Haan, D., Goodson, A.D., Staal, C., Vaughan, N.The biology of the harbour porpoise (Eds. Read, A.J., Wiepkema, P.R. and Nachtigall, P.E.). De Spil Publishers, Woerden, The Netherlands, 367-383Acoustics / Behavioral response studies (Effect of sound on behavior)
2000"The effects of acoustic alarms on the behavior of the harbor porpoises (phocoena phocoena) in a floating pen" [DOI]Kastelein, R.A., Rippe, T., Vaughan, N., Schooneman, N.M., Verboom, W.C., de Haan, D.Marine Mammal Science 16, 46-64Acoustics / Behavioral response studies (Effect of sound on behavior)
2001"The influence of three acoustic alarms on the behaviour of harbour porpoises (Phocoena phocoena) in a floating pen" [DOI]Kastelein, R.A., de Haan, D., Vaughan, N., Staal, C., Schooneman, N.M.Marine Environmental Research 52, 351-371Acoustics / Behavioral response studies (Effect of sound on behavior)
2005"The influence of acoustic emissions for underwater data transmission on the behaviour of harbour porpoises (Phocoena phocoena) in a floating pen" [DOI]Kastelein, R.A., Verboom, W.C., Muijsers, M., Jennings, N.V., van der Heul, S.Marine Environmental Research 59, 287-307Acoustics / Behavioral response studies (Effect of sound on behavior)
2006"The influence of underwater data transmission sounds on the displacement of captive harbour seals (Phoca vitulina)"Kastelein, R.A., van der Heul, S., Verboom, W.C., Triesscheijn, R.J.V., Vaughan- Jennings, N.Marine Environmental Research 61, 19-39Acoustics / Behavioral response studies (Effect of sound on behavior)
2006"Differences in the response of a striped dolphin (Stenella coeruleoalba) and a harbour porpoise (Phocoena phocoena) to an acoustic alarm" [DOI]Kastelein, R.A., Jennings, N., Verboom, W.C., de Haan, D., Schooneman, N.M.Marine Environmental Research 61, 363-378Acoustics / Behavioral response studies (Effect of sound on behavior)
2006"Deterring effects of 8-45 kHz tone pulses on harbor seals (Phoca vitulina) in a large pool"Kastelein, R.A., van der Heul, S, Terhune, J.M., Verboom, W.C., Triesscheijn, R.J.V.Marine Environmental Research 62, 356-373Acoustics / Behavioral response studies (Effect of sound on behavior)
2007"Effects of acoustic alarms, designed to reduce small cetacean bycatch, on the behaviour of North Sea fish species in a large tank"Kastelein, R.A., van der Heul, S, van der Veen, J., Verboom, W.C., Jennings, N., Reijnders P.Marine Environmental Research 64, 160-180Acoustics / Behavioral response studies (Effect of sound on behavior)
2008"Startle response of captive North Sea fish species to underwater tones between 0.1 and 64 kHz"Kastelein, R.A., van der Heul, S, Verboom, W.C., Jennings, N., van der Veen, J., de Haan, D.Marine Environmental Research 65, 369-377Acoustics / Behavioral response studies (Effect of sound on behavior)
2008"Behavioral avoidance threshold level of a harbor porpoise (Phocoena phocoena) for a continuous 50 kHz pure tone (L)"Kastelein, R.A., Verboom, W.C., Jennings, N., de Haan, D.J. Acoust. Soc. Am. 123, 1858-1861. DOI: 10.1121/1.2874557Acoustics / Behavioral response studies (Effect of sound on behavior)
2008"Effect of vibrations on the behavior of cockles (bivalve molluscs)"Kastelein, R.A.Bioacoustics 17, 74-75 (abstract)Acoustics / Behavioral response studies (Effect of sound on behavior)
2008"Acoustic dose-effect relationships in marine fish"Kastelein, R.A., van der Heul, S., Verboom, W.C., de Haan, D., Reijnders, P.Bioacoustics 17, 201-202 (abstract)Acoustics / Behavioral response studies (Effect of sound on behavior)
2008"The influence of 70 and 120 kHz tonal signals on the behavior of harbor porpoises (Phocoena phocoena) in a floating pen" [URL]Kastelein, R.A., Verboom, W.C., Jennings, N., de Haan, D., van der Heul, S.Marine Environmental Research 66, 319-326Acoustics / Behavioral response studies (Effect of sound on behavior)
2011"Effect of broadband-noise masking on the behavioral response of a harbor porpoise (Phocoena phocoena) to 1-s duration 6-7 kHz sonar up-sweeps" [URL]Kastelein, R.A., Steen, N., de Jong, C.A.F., Wensveen, P.J., Verboom, W.C.J. Acoust. Soc. Am. 129, 2307-2315Acoustics / Behavioral response studies (Effect of sound on behavior)
2012"Impacts of Anthropogenic Sounds on Phocoena phocoena (Harbor Porpoise)"Kastelein, R.A., Jennings, N.The effects of Noise on Aquatic Life. Eds. A.N. Popper and A. Hawkins). Advances in Experimental Medicine and Biology, Springer Science +Business Media, 311-315Acoustics / Behavioral response studies (Effect of sound on behavior)
2012"Threshold received sound pressure levels of single 1-2 kHz and 6-7 kHz up-sweeps and down-sweeps causing startle responses in a harbor porpoise (Phocoena phocoena)" [URL]Kastelein, R.A., Steen, N., Gransier, R., Wensveen, P.J., de Jong, C.A.F.J. Acoust. Soc. Am. 131, 2325-2333Acoustics / Behavioral response studies (Effect of sound on behavior)
2013"Brief behavioral response threshold levels of a harbor porpoise (Phocoena phocoena) to five helicopter dipping sonar signals (1.33 to 1.43 kHz)"Kastelein, R.A., Gransier, R., van den Hoogen, Marjan, Hoek, L.Aquatic Mammals 39, 162-173, DOI 10.1578/AM.39.2.2013.162Acoustics / Behavioral response studies (Effect of sound on behavior)
2013"Behavioral responses of a harbor porpoise (Phocoena phocoena) to playbacks of broadband pile driving sounds"Kastelein, R.A., van Heerden, D., Gransier, R., Hoek, L.Marine Environmental Research 92, 206-214, DOI: 10.1016/j.marenvres.2013.09.020Acoustics / Behavioral response studies (Effect of sound on behavior)
2013"Brief behavioral response threshold level of a harbor porpoise (Phocoena phocoena) to an impulsive sound"Kastelein, R.A., Steen, N., Gransier, R., de Jong, C.A.F.Aquatic Mammals 39, 315-323, DOI 10.1578/AM.39.4.2013.315Acoustics / Behavioral response studies (Effect of sound on behavior)
2014"Effect of series of 1-2 kHz and 6-7 kHz up-sweeps and down-sweeps on the behavior of a harbor porpoise (Phocoena phocoena)"Kastelein, R.A., Schop, J., Gransier, R., Steen, N., Jennings, N.Aquatic Mammals 40(3), 232-242, DOI 10.1578/AM.40.3.2014.232Acoustics / Behavioral response studies (Effect of sound on behavior)
2014"Temporal structure of sound affects behavioural recovery from noise impact in European seabass"Neo, Y.Y., Seitz, J., Kastelein, R.A., Winter, H.V., ten Cate, C., Slabbekoorn, H.Biological Conservation 178, 65–73Acoustics / Behavioral response studies (Effect of sound on behavior)
2015"Hearing thresholds of harbor seals (Phoca vitulina) for playbacks of seal scarer signals, and effects of the signals on behavior"Kastelein, R.A., Helder-Hoek, L., Gransier, R., Terhune, J.M., Jennings, N., de Jong, C.A.F.Hydrobiologia 756, 75-88. DOI: 10.1007/s10750-014-2152-6Acoustics / Behavioral response studies (Effect of sound on behavior)
2015"Behavioral responses of a harbor porpoise (Phocoena phocoena) to 25-kHz FM Sonar Signals"Kastelein, R.A., van den Belt, I., Helder-Hoek, L., Gransier, R., Johansson, T.Aquatic Mammals 41, 311-326. DOI 10.1578/AM.41.3.2015.311Acoustics / Behavioral response studies (Effect of sound on behavior)
2015"Behavioral responses of harbor seals (Phoca vitulina) to sonar signals in the 25-kHz Range"Kastelein, R.A., Helder-Hoek, L., Janssens, G., Gransier, R., Johansson, T.Aquatic Mammals 41, 388-399. DOI 10.1578/AM.41.4.2015.388Acoustics / Behavioral response studies (Effect of sound on behavior)
2015"Behavioral responses of a harbor porpoise (Phocoena phocoena) to 24.5-25.5 kHz sonar down-sweeps with and without side bands"Kastelein, R.A., van den Belt, I., Gransier, R., Johansson, T.Aquatic Mammals 41, 400-411. DOI 10.1578/AM.41.4.2015.400Acoustics / Behavioral response studies (Effect of sound on behavior)
2015"Hearing thresholds of a harbor porpoise (Phocoena phocoena) for playbacks of seal scarer signals, and effects of the signals on behavior"Kastelein, R.A., Hoek, L., Gransier, R., de Jong, C.A.F., Terhune, J.M., Jennings, N.Hydrobiologia 756, 89-103. DOI 10.1007/s10750-014-2035-xAcoustics / Behavioral response studies (Effect of sound on behavior)
2016"Noise Impact on European Sea Bass Behavior: Temporal Structure Matters"Neo, Y. Y., Seitz, J., Kastelein, R.A., Winter, H.V., ten Cate, C., Slabbekoorn, H.The Effects of Noise on Aquatic Life II, Advances in Experimental Medicine and Biology 875, Springer Science+Business Media New York 2016 763 (Eds. A.N. Popper, A. Hawkins), 763-766. DOI 10.1007/978-1-4939-2981-8_93Acoustics / Behavioral response studies (Effect of sound on behavior)
2017"Behavioral responses of a harbor porpoise (Phocoena phocoena) to sounds from an Acoustic Porpoise Deterrent"Kastelein, R.A., Huybrechts, J., Covi, J., Helder-Hoek, L.Aquatic Mammals, 43, 233-244, DOI 10.1578/AM.43.3.2017.233Acoustics / Behavioral response studies (Effect of sound on behavior)
2017"Behavioral responses of harbor seals (Phoca vitulina) to FaunaGuard Seal Module sounds at two background noise levels"Kastelein, R.A., Horvers, M., Helder-Hoek, L., Van de Voorde, S., ter Hofstede, R., van der Meij, H.Aquatic Mammals 43, 347-363, DOI 10.1578/AM.43.4.2017.347Acoustics / Behavioral response studies (Effect of sound on behavior)
2017"Acoustic dose-behavioral response relationship in sea bass (Dicentrarchus labrax) exposed to playbacks of pile driving sounds"Kastelein, R.A., Jennings, N., Kommeren, A., Helder-Hoek, L., Schop, J.Marine Environmental Research 130, 315-324 DOI: 10.1016/j.marenvres.2017.08.010Acoustics / Behavioral response studies (Effect of sound on behavior)
2018"Spatial avoidance to experimental increase of intermittent and continuous sound in two captive harbour porpoises" [URL]Kok, A.C.M., Pamela Engelberts, J.P., Kastelein, R.A., Helder-Hoek, L., Van de Voorde, S., Visser, F., Slabbekoorn, H.Environmental Pollution 233 (2018) 1024e1036Acoustics / Behavioral response studies (Effect of sound on behavior)
2018"Swimming speed of a harbor porpoise (Phocoena phocoena) during playbacks of pile driving sounds"Kastelein, R.A., Van de Voorde, S., Jennings, N.Aquatic Mammals, 44, 92-99, DOI 10.1578/AM.44.1.2018.92Acoustics / Behavioral response studies (Effect of sound on behavior)
2018"Behavioral responses of harbor porpoises (Phocoena phocoena) to sonar playback sequences of sweeps and tones (3.5-4.1 kHz)"Kastelein, R.A., Helder-Hoek, L., Van de Voorde, S., de Winter, S., Janssen, S., Ainslie, M.A.Aquatic Mammals 44(4), 402-417.DOI 10.1578/AM.44.4.2018.389Acoustics / Behavioral response studies (Effect of sound on behavior)
2019"Behavioral Responses of Harbor Porpoises (Phocoena phocoena) to U.S. Navy 53C Sonar Signals in Noise"Kastelein, R.A., Ainslie, M.A., van Kester, R.Aquatic Mammals, 45(4), 359-366, DOI 10.1578/AM.45.4.2019.359Acoustics / Behavioral response studies (Effect of sound on behavior)
2019"Effect of pile-driving playback sound level on fish-catching efficiency in harbor porpoises (Phocoena phocoena)"Kastelein, R.A., Huijser, L. A. E., Cornelisse, S., Helder-Hoek, L., Jennings, N., de Jong, C.A.F.Aquatic Mammals 2019, 45(4), 398-410, DOI 10.1578/AM.45.4.2019.398Acoustics / Behavioral response studies (Effect of sound on behavior)
2019"Behavioral responses of a harbor porpoise (Phocoena phocoena) to a series of four different simulated low-frequency sonar sounds (1.33-1.43 kHz)"Kastelein, R.A., Verhoeven, A., Helder-Hoek, L.Aquatic Mammals, 45(6), 632-645, DOI 10.1578/AM.45.6.2019.632Acoustics / Behavioral response studies (Effect of sound on behavior)
2019"Effect of a bubble screen on the behavioral responses of captive harbor porpoises (Phocoena phocoena) exposed to airgun sounds"Kastelein, R.A., von Benda-Beckmann, A.M., Lam, F-P. A., Jansen, E., de Jong, C.A.F.Aquatic Mammals 45(6), 706-716, DOI 10.1578/AM.45.6.2019.706Acoustics / Behavioral response studies (Effect of sound on behavior)
2022"Behavioral responses of a harbor porpoise (Phocoena phocoena) depend on the frequency content of pile-driving sounds"Kastelein, R.A., De Jong, C.A.F., Tougaard, J., Helder-Hoek, L., Defillet, L.N.Aquatic Mammals, 48(2), 97-109, DOI 10.1578/AM.48.2.2022.97Acoustics / Behavioral response studies (Effect of sound on behavior)
2023"Atlantic Green Turtles and Hawksbill Turtles: Behavioral Responses to Sound." [URL]Kastelein, R.A., Smink, A., Jennings, N.The Effects of Noise on Aquatic Life, Springer Nature Switzerland AG (A. N. Popper et al., eds.)Acoustics / Behavioral response studies (Effect of sound on behavior)
2025"Effect of impact pile-driving playback sound on harbor seal (Phoca vitulina) behavior: dose-response relationship and frequency weighting" [DOI]Kastelein, R.A., de Jong, C., Helder-Hoek, L., Defillet, L., Terhune, J.M., Kuiphof, F., Van Acoleyen, L.Aquatic Mammals 51(4), 319-335Acoustics / Behavioral response studies (Effect of sound on behavior)
2026"Reducing Fisheries Bycatch: Behavioral Responses of a Harbor Porpoise (Phocoena phocoena) in a Pool to the Sounds of a 'Banana Pinger'" [DOI]Kastelein, R.A., Hoek, L., Siemensma, M.L., Therhune, J.M., Jennings, M.A. N. Popper et al. (eds.), The Effects of Noise on Aquatic Life IVAcoustics / Behavioral response studies (Effect of sound on behavior)
1988"Low-frequency underwater hearing sensitivity in belugas, Delphinapterus leucas"Awbrey, F.T., Thomas, J.A., Kastelein, R.A.J. Acoust. Soc. Am. 84, 2273-2275Acoustics / Hearing
1993"Free field hearing tests on wild Atlantic walruses (Odobenus rosmarus rosmarus) in air"Kastelein, R.A., Van Ligtenberg, C.L., Gjertz, I. and Verboom, W.C.Aquatic Mammals 19, 143-148Acoustics / Hearing
1996"Aerial hearing sensitivity tests with a male Pacific walrus (Odobenus rosmarus divergens), in the free field and with headphones"Kastelein, R.A., Mosterd, P., Ligtenberg, C.L., van and Verboom, W.C.Aquatic Mammals 22, 81-93Acoustics / Hearing
1997"Detection of bone conductor signals by a harbour porpoise (Phocoena phocoena): a pilot study"Kastelein, R.A., Kaay, L.S.P. van der, Staal, C., Schooneman, N.M., Ligtenberg, C.L. vanThe biology of the harbour porpoise (Eds. Read, A.J., Wiepkema, P.R. and Nachtigall, P.E.). De Spil Publishers, Woerden, The Netherlands, 313-327Acoustics / Hearing
1997"Low-frequency aerial hearing of a harbour porpoise (Phocoena phocoena)"Kastelein, R.A., Nieuwstraten, S.H., Staal, C., Ligtenberg, C.L. van, Versteegh, D.The biology of the harbour porpoise (Eds. Read, A.J., Wiepkema, P.R. and Nachtigall, P.E.). De Spil Publishers, Woerden, The Netherlands, 295-312Acoustics / Hearing
2002"Audiogram of a harbor porpoise (Phocoena phocoena) measured with narrow-band frequency-modulated signals" [DOI]Kastelein, R.A., Bunskoek, P., Hagedoorn, M., Au, W.W.L., de Haan, D.J. Acoust. Soc. Am. 112, 334-344Acoustics / Hearing
2002"Underwater audiogram of a Pacific walrus (Odobenus rosmarus divergens) measured with narrow-band frequency-modulated signals" [DOI]Kastelein, R.A., Mosterd, P., van Santen, B., Hagedoorn, M., de Haan, D.J. Acoust. Soc. Am. 112, 2173-2182Acoustics / Hearing
2003"Audiogram of a striped dolphin (Stenella coeruleoalba)" [DOI]Kastelein, R.A., Hagedoorn, M., Au, W.W.L., de Haan, D.Journal of the Acoustical Society of America 113(2), 1130-1137Acoustics / Hearing
2005"Receiving beam patterns in the horizontal plane of a harbor porpoise (Phocoena phocoena)" [DOI]Kastelein, R.A., Janssen, M., Verboom, W.C., de Haan, D.J. Acoust. Soc. Am. 118, 1172-1179Acoustics / Hearing
2005"Underwater hearing sensitivity of a male and a female Steller sea lion (Eumetopias jubatus)" [DOI]Kastelein, R.A., van Schie, R., Verboom, W.C., C and de Haan, D.J. Acoust. Soc. Am. 118, 1820-1829Acoustics / Hearing
2007"The influence of signal parameters on the sound source localization ability of a harbor porpoise (Phocoena phocoena)" [DOI]Kastelein, R.A., de Haan, D., Verboom, W.C.J. Acoust. Soc. Am. 122, 1238-1248Acoustics / Hearing
2008"Effect of two levels of masking noise on the hearing threshold of a harbor porpoise (Phocoena phocoena) for a 4.0 kHz signal"Kastelein, R.A., Wensveen, P.J.Aquatic Mammals 34, 420-425. DOI 10.1578/AM.34.4.2008.420Acoustics / Hearing
2009"Underwater detection of tonal signals between 0.125 and 100 kHz by harbor seals (Phoca vitulina)"Kastelein, R.A., Wensveen, P.J., Hoek, L., Verboom, W.C., Terhune, J.M.J. Acoust. Soc. Am. 125, 1222-1229. DOI: 10.1121/1.3050283Acoustics / Hearing
2009"Underwater hearing sensitivity of harbor seals (Phoca vitulina) for narrow noise bands between 0.2 and 80 kHz"Kastelein, R.A., Wensveen, P.J., Hoek, L., Verboom, W.C., Terhune, J.M.J. Acoust. Soc. Am. 126, 476-483. DOI: 10.1121/1.3132522Acoustics / Hearing
2009"Critical ratios in harbor porpoises (Phocoena phocoena) for tonal signals between 0.315 and 150 kHz in random Gaussian white noise" [URL]Kastelein, R.A., Wensveen, P.J., Hoek, L., Au, W.W.L., Terhune, J.M., de Jong, C.A.F.J. Acoust. Soc. Am. 126, 1588-1597Acoustics / Hearing
2010"The effect of signal duration on the underwater detection thresholds of a harbor porpoise (Phocoena phocoena) for single frequency-modulated tonal signals between 0.25 and 160 kHz" [URL]Kastelein, R.A., Hoek, L., de Jong, C.A.F., Wensveen, P.J.J. Acoust. Soc. Am. 128, 3211-3222Acoustics / Hearing
2010"The effect of signal duration on the underwater hearing thresholds of two harbor seals (Phoca vitulina) for single tonal signals between 0.2 and 40 kHz"Kastelein, R.A., Hoek, L., Wensveen, P.J., Terhune, J.M., de Jong, C.A.F.JASA, 127, 1135-1145. DOI: 10.1121/1.3283019Acoustics / Hearing
2011"Hearing thresholds of a harbor porpoise (Phocoena phocoena) for sweeps (1-2 kHz and 6-7 kHz bands) mimicking naval sonar signals" [URL]Kastelein, R.A., Hoek, L., de Jong, C.A.F.J. Acoust. Soc. Am. 129, 3393-3399Acoustics / Hearing
2011"Hearing thresholds of a harbor porpoise (Phocoena phocoena) for helicopter dipping sonar signals (1.43-1.33 kHz) (L)"Kastelein, R.A., Hoek, L., de Jong, C.A.F.J. Acoust. Soc. Am. 130, 679-682. DOI: 10.1121/1.3605541Acoustics / Hearing
2011"Near-threshold equal-loudness contours for harbor seals (Phoca vitulina) derived from reaction times during underwater audiometry: a preliminary study"Kastelein, R.A., Wensveen, P.J., Terhune, J.M., de Jong, C.A.F.J. Acoust. Soc. Am. 129, 488-495. DOI: 10.1121/1.3518779Acoustics / Hearing
2012"The hearing threshold of a harbor porpoise (Phocoena phocoena) for impulsive sounds (L)" [URL]Kastelein, R.A., Gransier, R., Hoek, L., de Jong, C.A.F.J. Acoust. Soc. Am. 132, 607-610Acoustics / Hearing
2012"Hearing threshold shifts and recovery in harbor seals (Phoca vitulina) after octave-band noise exposure at 4 kHz" [URL]Kastelein, R.A., Gransier, R., Hoek, L., Macleod, A., Terhune, J.M.J. Acoust. Soc. Am. 132, 2745-2761Acoustics / Hearing
2012"Temporary threshold shifts and recovery in a harbor porpoise (Phocoena phocoena) after octave-band noise at 4 kHz" [URL]Kastelein, R.A., Gransier, R., Hoek, L., Olthuis, J.J. Acoust. Soc. Am. 132, 3525-3537Acoustics / Hearing
2013"Hearing frequency thresholds of a harbor porpoise (Phocoena phocoena) temporarily affected by a continuous 1.5 kHz tone" [URL]Kastelein, R.A., Gransier, R., Hoek, L., Rambags, M.J. Acoust. Soc. Am. 134, 2286-2292Acoustics / Hearing
2013"Comparative temporary threshold shifts in a harbor porpoise and harbor seal, and severe shift in a seal (L)" [URL]Kastelein, R.A., Gransier, R., Hoek, L.J. Acoust. Soc. Am. 134(1), 13-16Acoustics / Hearing
2013"Hearing thresholds of two harbor seals (Phoca vitulina) for playbacks of multiple pile driving strike sounds" [URL]Kastelein, R.A., Hoek, L., Gransier, R., de Jong, C.A.F., Jennings, N.J. Acoust. Soc. Am. 134, 2307-2312Acoustics / Hearing
2013"Hearing thresholds of a harbor porpoise (Phocoena phocoena) for playbacks of multiple pile driving strike sounds" [URL]Kastelein, R.A., Hoek, L., Gransier, R., de Jong, C.A.F.J. Acoust. Soc. Am. 134, 2302-2306Acoustics / Hearing
2014"Equal latency contours and auditory weighting functions for the harbour porpoise (Phocoena phocoena)"Wensveen, P.J., Huijser, L.A.E., Hoek, L., Kastelein, R.A.J. Exp. Biol. 217, 1-11 DOI:10.1242/jeb.091983Acoustics / Hearing
2014"Effect of level, duration, and inter-pulse interval of 1-2 kHz sonar signal exposures on harbor porpoise hearing" [URL]Kastelein, R.A., Hoek, L., Gransier, R., Rambags, M., Claeys, N.J. Acoust. Soc. Am. 136, 412-422Acoustics / Hearing
2014"Frequency of greatest temporary hearing threshold shift in harbor porpoises (Phocoena phocoena) depends on the noise level"Kastelein, R.A., Schop, J., Gransier, R., Hoek, L.J. Acoust. Soc. Am. 136, 1410-1418, DOI: 10.1121/1.4892794Acoustics / Hearing
2015"Hearing thresholds of harbor seals (Phoca vitulina) for playbacks of seal scarer signals, and effects of the signals on behavior"Kastelein, R.A., Helder-Hoek, L., Gransier, R., Terhune, J.M., Jennings, N., de Jong, C.A.F.Hydrobiologia 756, 89-103. DOI 10.1007/s10750-014-2035-xAcoustics / Hearing
2015"Hearing frequencies of a harbor porpoise (Phocoena phocoena) temporarily affected by played back offshore pile driving sounds"Kastelein, R.A., Gransier, R., Marijt, M. A.T., Hoek, L.J. Acoust. Soc. Am. 137, 556–564, DOI: 10.1121/1.4906261Acoustics / Hearing
2015"Effect of intermittent and continuous 6-7 kHz sonar sweep exposures on harbor porpoise (Phocoena phocoena) hearing"Kastelein, R.A., Gransier, R., Schop, J., Hoek, L.J. Acoust. Soc. Am. 137, 1623-1633. DOI: 10.1121/1.4916590Acoustics / Hearing
2015"Hearing thresholds of a harbor porpoise (Phocoena phocoena) for playbacks of seal scarer signals, and effects of the signals on behavior"Kastelein, R.A., Hoek, L., Gransier, R., de Jong, C.A.F., Terhune, J.M., Jennings, N.Hydrobiologia 756, 89-103. DOI 10.1007/s10750-014-2035-xAcoustics / Hearing
2015"Assessing the Impact of Underwater Clearance of Unexploded Ordnance on Harbour Porpoises (Phocoena phocoena) in the Southern North Sea"von Benda-Beckmann, A.M., Aarts, G., Sertlek, H.O., Lucke, C., Verboom, W.C., Kastelein, R.A., Ketten, D.R., van Bemmelen, R., Lam, F-P, Kirkwood, R.J., Ainslie, M.A.Aquatic Mammals 41, 503-523. DOI 10.1578/AM.41.4.2015.503Acoustics / Hearing
2015"Hearing thresholds of a harbor porpoise (Phocoena phocoena) for narrow-band sweeps" [URL]Kastelein, R.A., Schop, J., Hoek, L., Covi, J.J. Acoust. Soc. Am. 138, 2508–2512Acoustics / Hearing
2016"Conditioned hearing sensitivity change in the harbor porpoise (Phocoena phocoena)"Nachtigall, P.E., Supin, A. Ya. Pacini, A.F., Kastelein, R.A.J. Acoust. Soc.Am. 140, 960–967. DOI: 10.1121/1.4960783Acoustics / Hearing
2016"Pile driving playback sounds and temporary threshold shift in harbor porpoises (Phocoena phocoena): Effect of exposure duration" [URL]Kastelein, R.A., Helder-Hoek, L., Covi, J., Gransier, R.J. Acoust. Soc. Am. Am. 139, 2842-2851Acoustics / Hearing
2016"The Cumulative Effects of Exposure to Continuous and Intermittent Sounds on Temporary Hearing Threshold Shifts Induced in a Harbor Porpoise (Phocoena phocoena)"Kastelein, R.A., Hoek, L., Gransier, R.The Effects of Noise on Aquatic Life II, Advances in Experimental Medicine and Biology 875, Springer Science+Business Media New York 2016 763 (Eds. A.N. Popper, A. Hawkins), 523-528. DOI 10.1007/978-1-4939-2981-8_93Acoustics / Hearing
2017"Hearing thresholds of a male and a female harbor porpoise (Phocoena phocoena)" [URL]Kastelein, R.A., Helder-Hoek, L., Van de Voorde, S.J. Acoust. Soc. Am. 142, 1006–1010Acoustics / Hearing
2017"Effects of exposure to 53-C sonar playback sounds (3.5-4.1 kHz) on harbor porpoise (Phocoena phocoena) hearing" [URL]Kastelein, R.A., Helder-Hoek, L., Van de Voorde, S.J. Acoust. Soc. Am. 142, 1965–1975Acoustics / Hearing
2017"Temporary hearing threshold shift in a harbor porpoise (Phocoena phocoena) after exposure to multiple airgun sounds"Kastelein, R.A., Helder-Hoek, L., Van de Voorde, S., von Benda-Beckmann, A.M., Lam, F-P. A., Jansen, E., de Jong, C.A.F., Ainslie, M.A.J. Acoust. Soc. Am. 142, 2430-2442. DOI: 10.1121/1.5007720Acoustics / Hearing
2018"Four odontocete species change hearing levels when warned of impending loud sound"Nachtigall, P.E., Supin, A. Ya., Pacini, A.F., Kastelein, R.A.Integrative Zoology 13, 2–20. doi: 10.1111/1749-4877.12286Acoustics / Hearing
2018"Effect of pile driving sounds on harbor seals (Phoca vitulina) hearing"Kastelein, R.A., Hoek, L., Kommeren, A., Covi, J., Gransier, R.J. Acoust. Soc. 143, 3583–3594. DOI: 10.1121/1.5040493Acoustics / Hearing
2018"Hearing thresholds, for underwater sounds, of harbor seals (Phoca vitulina) at the water surface" [URL]Kastelein, R.A., Helder-Hoek, L., Terhune, J.M.J. Acoust. Soc. Am. 143, 2554-2563Acoustics / Hearing
2019"Temporary hearing threshold shift in harbor porpoises (Phocoena phocoena) due to one-sixth-octave noise band at 32 kHz"Kastelein, R.A., Helder-Hoek, L., Cornelisse, S.A., Huijser, L.A.E., Gransier, R.Aquatic Mammals, 45(5), 549-562, DOI 10.1578/AM.45.5.2019.549Acoustics / Hearing
2019"Hearing thresholds of two harbor seals (Phoca vitulina) for helicopter dipping sonar signals (1.3-1.4 kHz)"Kastelein, R.A., Gransier, R., Brouwers, M., Helder-Hoek, L.Aquatic Mammals, 45(3), 349-355, DOI 10.1578/AM.45.3.2019.349Acoustics / Hearing
2019"Frequency of greatest temporary threshold shift in harbor seals (Phoca vitulina) depends on the fatiguing sound level" [URL]Kastelein, R.A., Helder-Hoek, L., Gransier, R.J. Acoust. Soc. Am. 145, 1353-1362Acoustics / Hearing
2019"Temporary hearing threshold shift in harbor seals (Phoca vitulina) due to a one-sixth-octave noise band centered at 16 kHz"Kastelein, R.A., Helder-Hoek, L., Cornelisse, S.A., Huijser, L.A.E., Terhune, J.M.J. Acoust. Soc. Am. 146, 3113–3122, DOI 10.1121/1.5130385Acoustics / Hearing
2020"Temporary hearing threshold shift in harbor porpoises (Phocoena phocoena) due to one-sixth-octave noise band at 16 kHz"Kastelein, R.A., Helder-Hoek, L., van Kester, R., Huisman, R., Gransier, R.Aquatic Mammals 45, 280-292, DOI 10.1578/AM.45.3.2019.280Acoustics / Hearing
2020"Temporary hearing threshold shift at ecologically relevant frequencies in a harbor porpoise (Phocoena phocoena) due to exposure to a noise band centered at 88.4 kHz"Kastelein, R.A., Helder-Hoek, L., Cornelisse, S.A., Huijser, L.A.E., Gransier, R.Aquatic Mammals, 46(5), 444-453, DOI 10.1578/AM.46.5.2020.444Acoustics / Hearing
2020"Temporary threshold shift in a second harbor porpoise (Phocoena phocoena) after exposure to a noise band at 1.5 kHz and a 6.5 kHz continuous wave"Kastelein, R.A., Helder-Hoek, L., Cornelisse, S.A., Defillet, L.N., Huijser, L.A.E.Aquatic Mammals, 46(5), 431-443, DOI 10.1578/AM.46.5.2020.431Acoustics / Hearing
2020"Temporary hearing threshold shift in harbor seals (Phoca vitulina) due to one-sixth-octave noise bands centered at 0.5, 1, and 2 kHz"Kastelein, R.A., Helder-Hoek, L., Cornelisse, S.A., Defillet, L. N., Huijser, L.A.E., Terhune, J.M.J. Acoust. Soc. Am. 148 (6), 3873-3885, DOI 10.1121/10.0002781Acoustics / Hearing
2020"Temporary hearing threshold shift in harbor seals (Phoca vitulina) due to a one-sixth-octave noise band centered at 40 kHz"Kastelein, R.A., Parlog, C., Helder-Hoek, L., Cornelisse, S.A., Huijser, L.A.E., Terhune, J.M.J. Acoust. Soc. Am. 147 (3), 1966-1976. DOI 10.1121/10.0000908Acoustics / Hearing
2020"Temporary hearing threshold shift in harbor seals (Phoca vitulina) due to a one-sixth-octave noise band centered at 32 kHz"Kastelein, R.A., Helder-Hoek, L., Cornelisse, S.A., Huijser, L.A.E., Terhune, J.M.J. Acoust. Soc. Am. 147 (3), 1885-1896. DOI 10.1121/10.0000889Acoustics / Hearing
2020"Temporary hearing threshold shift in harbor porpoises (Phocoena phocoena) due to one-sixth-octave noise bands at 63 kHz"Kastelein, R.A., Cornelisse, S.A., Huijser, L.A.E., Helder-Hoek, L.Aquatic Mammals 46(2), 167-182, DOI 10.1578/AM.46.2.2020.167Acoustics / Hearing
2020"Lack of reproducibility of temporary hearing threshold shifts in a harbor porpoise after exposure to repeated airgun sounds" [URL]Kastelein, R.A., Helder-Hoek, L., Cornelisse, S.A., von Benda-Beckmann, A.M., Lam, F-P. A., de Jong, C.A.F., Ketten, D.R.J. Acoust. Soc. Am. 148 (2), 556–565Acoustics / Hearing
2021"Masking release at 4 kHz in harbor porpoises (Phocoena phocoena) associated with sinusoidal amplitude-modulated masking noise"Kastelein, R.A., Helder-Hoek, L., Covi, J., Terhune, J.M., Klump, G.The Journal of the Acoustical Society of America 150, 1721-1732, doi: 10.1121/10.0006103Acoustics / Hearing
2021"Temporary hearing threshold shift in California sea lions (Zalophus californianus) due to one-sixth-octave noise bands centered at 2 and 4 kHz: effect of duty cycle and testing the equal-energy hypothesis"Kastelein, R.A., Helder-Hoek, L., Defillet, L. N., Huijser, L.A.E., Terhune, J.M., Gransier, R.Aquatic Mammals, 47(4), 394-418, DOI 10.1578/AM.47.4.2021.394Acoustics / Hearing
2021"Temporary hearing threshold shift in a harbor porpoise (Phocoena phocoena) due to exposure to a continuous one-sixth-octave noise band centered at 0.5 kHz"Kastelein, R.A., Helder-Hoek, L., Cornelisse, S.A., Defillet, L. N., Huijser, L.A.E., Gransier, R.Aquatic Mammals, 47(2) 135-145, DOI 10.1578/AM.47.2.2021.135Acoustics / Hearing
2022"Evaluation of kurtosis-corrected sound exposure level as a metric for predicting onset of hearing threshold shifts in harbor porpoises (Phocoena phocoena)"von Benda-Beckmann, A.M., Ketten, D. R., Lam, F. P. A., de Jong, C. A. F., Muller, R. A. J., Kastelein, R.A.J. Acoust. Soc. Am. 152 (1), 295–301. https://doi.org/10.1121/10.0012364Acoustics / Hearing
2022"Temporary hearing threshold shift in California sea lions (Zalophus californianus) due to one-sixth-octave noise bands centered at 0.6 and 1 kHz"Kastelein, R.A., Helder-Hoek, L., L., Defillet, L.N. Van Acoleyen, L. Huijser, L.A.E., Terhune, J.M.Aquatic Mammals 48(3), 248-265, DOI 10.1578/AM.48.3.2022.248Acoustics / Hearing
2022"Temporary hearing threshold shift in California sea lions (Zalophus californianus) due to one-sixth-octave noise bands centered at 8 and 16 kHz: Effect of duty cycle and testing the equal-energy hypothesis"Kastelein, R.A., Helder-Hoek, L., Defillet, L.N., Kuiphof, F., Huijser, L.A.E., Terhune, J.M.Aquatic Mammals, 48(1), 36-58, DOI 10.1578/AM.48.1.2022.36Acoustics / Hearing
2023"Underwater sound detection thresholds (0.031-80 kHz) of two California sea lions (Zalophus californianus) and a revised generic audiogram for the species" [DOI]Kastelein, R.A., Helder-Hoek, L., Van Acoleyen, L., Defillet, L.N., Huijser, L.A.E., Terhune, J.M.Aquatic Mammals 49(5), 422-435Acoustics / Hearing
2023"Masking release at 4 and 32 kHz in harbor seals associated with sinusoidal amplitude-modulated masking noise" [URL]Kastelein, R.A., Helder-Hoek, L., Defillet, L.N., Terhune, J. M., Beutelmann, R., Klump, G. M.J. Acoust. Soc. Am., 154 (1), 81–94Acoustics / Hearing
2024"Temporary hearing threshold shift and testing the equal-energy hypothesis in a harbor porpoise (Phocoena phocoena) after exposure to a continuous noise band at 8 kHz, and a revised TTS-onset function" [DOI]Kastelein, R.A., Helder-Hoek, L., Van Acoleyen, L., Defillet, L.N., Terhune, J.M.Aquatic Mammals, 50(5), 445-459Acoustics / Hearing
2024"Quantifying sound exposure in a pool: comparing hydrophones on a grid with a sound recording tag on a California sea lion (Zalophus californianus)" [DOI]Kastelein, R.A., Van Acoleyen, L., Tuytens, K., Jansen, E., Terhune, J.M., Jennings, N.Aquatic Mammals, 50(6), 592-610Acoustics / Hearing
2024"Susceptibility of harbor porpoise hearing to intermittent sound exposures. Springer Nature Switzerland AG. A. N. Popper et al. (eds.)" [DOI]von Benda-Beckmann, A.M., Kastelein, R.A., Lam, F-P. A., de Jong, C.A.F., Wensveen, P.J., Ketten, D.R.The Effects of Noise on Aquatic Life, 1-25Acoustics / Hearing
2024"Similar susceptibility to temporary hearing threshold shifts despite different audiograms in harbor porpoises and harbor seals" [DOI]Gransier, R., Kastelein, R.A.J. Acoust. Soc. Am. 155, 396–404Acoustics / Hearing
2025"Critical ratio measurements can be used to assess effective quiet thresholds in harbor seals, harbor porpoises, and California sea lions" [DOI]Terhune, J.M., Kastelein, R.A.Aquatic Mammals, 51(5), 28-37Acoustics / Hearing
2025"Similar hearing thresholds at 8 kHz, but dissimilar susceptibility of hearing to damage by 8 kHz sounds in a harbor porpoise, a harbor seal, and a California sea lion" [DOI]Kastelein, R.A., Jennings, N., Terhune, J.M.Aquatic Mammals, 51(5), 1-7Acoustics / Hearing
2025"Temporary hearing threshold shift in California sea lions (Zalophus californianus) due to a noise band centered at 40 kHz and Comparison with shifts due to lower-frequency sounds" [DOI]Kastelein, R.A., Helder-Hoek, L., Van Acoleyen, L., Defillet, L.N., Terhune, J.M., Jennings, N.Aquatic Mammals, 51(1) 13-30Acoustics / Hearing
2025"Temporary hearing threshold shift and testing the equal-energy hypothesis in harbor seals (Phoca vitulina) after exposure to a one-sixth-octave noise band centered at 8 kHz" [DOI]Kastelein, R.A., Helder-Hoek, L., Van Acoleyen, L., Terhune, J.M.Aquatic Mammals, 51(1) 79-95Acoustics / Hearing
1995"Echolocation signals of Harbour porpoises (Phocoena phocoena) in light and complete darkness"Kastelein, R.A., Nieuwstraten, S.H., Verboom, W.C.Harbour porpoises, laboratory studies to reduce bycatch (Eds Nachtigall, P.E., Lien, J., Au, W.W.L. and Read, A.J.). De Spil Publishers, Woerden, The Netherlands, 55-67Acoustics / Echolocation
1995"Source Levels and echolocation signal characteristics of juvenile Harbour porpoises (Phocoena phocoena) in a pool"Goodson, A.D., Kastelein, R.A., Sturtivant, C.R.Harbour porpoises, laboratory studies to reduce bycatch (Eds Nachtigall, P.E., Lien, J., Au, W.W.L. and Read, A.J.). De Spil Publishers, Woerden, The Netherlands, 41-53Acoustics / Echolocation
1995"Acoustic signals by Harbour porpoises (Phocoena phocoena)"Verboom, W.C., Kastelein, R.A.Harbour porpoises, laboratory studies to reduce bycatch (Eds Nachtigall, P.E., Lien, J., Au, W.W.L. and Read, A.J.). De Spil Publishers, Woerden, The Netherlands, 1-39Acoustics / Echolocation
1997"The ability of a harbour porpoise (Phocoena phocoena) to discriminate between objects buried in sand"Kastelein, R.A., Schooneman, N.M., Au, W.W.L., Verboom, W.C., Vaughan, N.The biology of the harbour porpoise (Eds. Read, A.J., Wiepkema, P.R. and Nachtigall, P.E.). De Spil Publishers, Woerden, The Netherlands, 329-342Acoustics / Echolocation
1997"Structure of harbour porpoise (Phocoena phocoena) click train signals"Verboom, W.C., Kastelein, R.A.The biology of the harbour porpoise (Eds. Read, A.J., Wiepkema, P.R. and Nachtigall, P.E.). De Spil Publishers, Woerden, The Netherlands, 343-362Acoustics / Echolocation
1999"Target detection by an echolocating harbor porpoise (Phocoena phocoena)"Kastelein, R.A., Au, W.W.L., Rippe, H.T., Schooneman, N.M.J. Acoust. Soc. Am. 105, 2493-2498.Acoustics / Echolocation
1999"Transmission beam pattern and echolocation signals of a harbor porpoise (Phocoena phocoena)"Au, W.W.L., Kastelein, R.A., Rippe, H.T., Schooneman, N.M.J. Acoust. Soc. Am. 106, 3699-3705Acoustics / Echolocation
2000"Detection distances of bottom-set gillnets by harbor porpoises (Phocoena phocoena) and bottlenose dolphins (Tursiops truncatus)"Kastelein, R.A., Au, W.W.L., de Haan, D.Marine Environmental Research 49, 359-375Acoustics / Echolocation
2002"Characteristics of echolocation signals used by a harbour porpoise (Phocoena phocoena) in a target detection experiment"Teilmann, J., Miller, L.A., Kirketerp, T., Kastelein, R.A., Madsen, P.T., Nielsen, B.K., Au, W.W.L.Aquatic Mammals, 28, 275-284Acoustics / Echolocation
2004"Structure of harbor porpoise (Phocoena phocoena) acoustic signals with high repetition rates"Verboom, W.C., Kastelein, R.A.Echolocation in Bats and Dolphins. Eds. J.A. Thomas, C.F. Moss and M. Vater. The University of Chicago Press, Chicago, 40-43Acoustics / Echolocation
2006"Testing the odontocete acoustic prey debilitation hypothesis: No stunning results" [DOI]Benoit-Bird, K.J., Au, W.W.L., Kastelein, R.A.J. Acoust. Soc. Am. 120, 1118-1123Acoustics / Echolocation
2006"Acoustic radiation from the head of echolocating harbor porpoises (Phocoena phocoena)" [DOI]Au, W.W.L., Kastelein, R.A., Benoit-Bird, K.J., Cranford, T.W., McKenna, M.F.J. of Exp. Biol. 209, 2726-2733Acoustics / Echolocation
2007"Modeling the detection range of fish by echolocating bottlenose dolphins and harbor porpoises" [DOI]Au, W.W.L., Benoit-Bird, K.J., Kastelein, R.A.J. Acoust. Soc. Am. 121, 3954-3962Acoustics / Echolocation
2008"Number and duration of echolocation click trains produced by a harbor porpoise (Phocoena phocoena) in relation to target and performance (L.)"Kastelein, R.A., Verlaan, M., Jennings, N.J. Acoust. Soc. Am. 124, 40-43. DOI: 10.1121/1.2924132Acoustics / Echolocation
2009"Acoustic basis for fish prey discrimination by echolocating dolphins and porpoises"Au, W.W.L., Branstetter, B.K., Benoit-Bird, K.J., Kastelein, R.A.J. Acoust. Soc. Am. 126, 460-467Acoustics / Echolocation
2019"Acoustic reflectivity of a harbor porpoise (Phocoena phocoena)" [URL]Au, W.W. L., Kastelein, R.A., Helder-Hoek, L.J. Acoust. Soc. Am. 146 (4), 2475–2481Acoustics / Echolocation
1995"Airborne vocalizations of Pacific walrus pups (Odobenus rosmarus divergens)" [DOI]Kastelein, R.A., Postma, J., Verboom, W.C.Aquatic Mammals, 48(1), 36-58Acoustics / Vocalizations
1995"Rutting whistles of a male Pacific walrus (Odobenus rosmarus divergens)"Verboom, W.C., Kastelein, R.A.Sensory Systems of Aquatic Mammals (Eds. Kastelein, R.A., Thomas, J.A. and Nachtigall, P.E.). De Spil Publishers, Woerden, The NetherlandsAcoustics / Vocalizations
1990"Marginal papillae on the tongue of Harbour porpoises (Phocoena phocoena), Bottlenose dolphins (Tursiops truncatus) and the Commerson's dolphin (Cephalorhynchus commersonii)"Kastelein, R.A., Dubbeldam, J.Aquatic Mammals 15,158-170Anatomy & morphology
1990"Anatomical and histological characteristics of the eyes of a month-old and adult Harbor porpoise (Phocoena phocoena)"Kastelein, R.A., Zweypfenning, R.C.V.J., Spekreijse, H.Sensory abilities of cetaceans/Laboratory and field evidence (Eds. J.A. Thomas and R.A. Kastelein) Plenum Press, New York,463-480Anatomy & morphology
1990"The relationship between body weight and morphological measurements in Harbour Porpoises (Phocoena phocoena) from the North Sea"Kastelein, R.A., van Battum, R.Aquatic Mammals 16, 48-52Anatomy & morphology
1990"The suckling of a Bottlenose dolphin calf (Tursiops truncatus) by a foster mother, and information on transverse birth bands"Kastelein, R.A., Dokter, T., Zwart, P.Aquatic Mammals 16,134-138Anatomy & morphology
1990"The anatomy of the Walrus head (Odobenus rosmarus). Part I: The skull"Kastelein, R.A., Gerrits, N. M.Aquatic Mammals 16,101-119Anatomy & morphology
1991"The anatomy of the Walrus head (Odobenus rosmarus) Part 2: Description of the muscles and of their role in feeding and haul-out behaviour"Kastelein, R.A., Gerrits, N. M., Dubbeldam, J. L.Aquatic Mammals 17,156-180Anatomy & morphology
1992"Foreign bodies in the stomach of a female Harbour porpoise (Phocoena phocoena) from the North Sea"Kastelein, R.A., Lavaleije, M.S.S.Aquatic Mammals 18, 40-46Anatomy & morphology
1993"The anatomy of the Walrus head (Odobenus rosmarus). Part 3: The eyes and their function in Walrus ecology"Kastelein, R.A., Zweipfenning, R.C.V.J., Spekreijse, H., Dubbeldam, J.L., Born, E.W.Aquatic Mammals 19, 61-92Anatomy & morphology
1996"The anatomy of the walrus head (Odobenus rosmarus). Part 4: The ears and their function in aerial and underwater hearing"Kastelein, R.A., Dubbeldam, J.L., de Bakker, M.A.G., Gerrits, N.M.Aquatic Mammals 22, 95-125Anatomy & morphology
1997"Blubber thickness in harbour porpoises (Phocoena phocoena)"Kastelein, R.A., Sijs, S.J. van der, Staal, C., Nieuwstraten, S.H.The biology of the harbour porpoise (Eds. Read, A.J., Wiepkema, P.R. and Nachtigall, P.E.). De Spil Publishers, Woerden, The Netherlands, 179-199Anatomy & morphology
1997"An anatomical atlas of an adult female harbour porpoise"Kastelein, R.A., Dubbeldam, J. L., Luksenburg, J., Staal, C., Immerseel, F.The biology of the harbour porpoise (Eds. Read, A.J., Wiepkema, P.R. and Nachtigall, P.E.). De Spil Publishers, Woerden, The Netherlands, 87-178Anatomy & morphology
1997"The anatomy of the walrus head (Odobenus rosmarus). Part 5: The tongue and its function in walrus ecology"Kastelein, R.A., Dubbeldam, J. L., de Bakker, M.A.G.Aquatic Mammals 23, 29-47Anatomy & morphology
1997"Histology of the grooved ventral pouch of the minke whale, Balaenoptera acutorostrata, with special reference to the occurrence of lamellated corpuscles"de Bakker, M.A.G., Kastelein, R.A., Dubbeldam, J. L.Can. J. Zool. 75, 563Anatomy & morphology
2009"A case study of congenital diaphragmatic hernia in a juvenile striped dolphin (Stenella coeruleoalba)"Kastelein, R.A., van Dooren, M. F., Tibboel, D.Aquatic Mammals 35, 32-35Anatomy & morphology
2016"Reversible bending of the dorsal fins of harbor porpoises (Phocoena phocoena) and a striped dolphin (Stenella coeruleoalba) in captivity"Kastelein, R.A., Triesscheijn, R.J.V., Jennings, N.Aquatic Mammals, 42(2),218-226, DOI 10.1578/AM.42.2.2016.218Anatomy & morphology
1988"Case study of the neonatal period of a Grey Seal pup (Halichoerus grypus) in captivity"Kastelein, R.A., Wiepkema, P.R.Aquatic Mammals, 14.1, 33-38Behavior
1990"The suckling period of a Grey seal (Halichoerus grypus) while confined to an outdoor land area"Kastelein, R.A., Wiepkema, P.R.Aquatic Mammals 16(3),120-128Behavior
1990"The suckling of a Bottlenose dolphin calf (Tursiops truncatus) by a foster mother, and information on transverse birth bands"Kastelein, R.A., Dokter, T., P. ZwartAquatic Mammals 16(3),134-138Behavior
1991"Swimming, diving, and respiration patterns of a Northern bottlenose whale (Hyperoodon ampullatus, Forster, 1770)"Kastelein, R.A., Gerrits, N.M.Aquatic Mammals, 17(1), 20-30Behavior
1991"The suckling period of a Grey seal (Halichoerus grypus) while the mother had access to a pool"Kastelein, R.A., Wiepkema, P.R., Vaughan, N.Aquatic Mammals, 17(1),42-51Behavior
1995"Behaviour of Harbour porpoises (Phocoena phocoena) in response to ropes. In: Harbour porpoises, laboratory studies to reduce bycatch (Eds Nachtigall, P.E., Lien, J., Au, W.W.L. and Read, A.J.)"Kastelein, R.A., de Haan, D., Staal, C.De Spil Publishers, Woerden, The Netherlands, 69-90Behavior
1995"Entanglement of Harbour porpoises (Phocoena phocoena) in fishing nets"Kastelein, R.A., de Haan, D., Staal, C., Nieuwstraten, S.H., Verboom, W.C.Harbour porpoises, laboratory studies to reduce bycatch (Eds Nachtigall, P.E., Lien, J., Au, W.W.L. and Read, A.J.). De Spil Publishers, Woerden, The Netherlands, 91-156Behavior
1995"Fishing technique development in a newly weaned Grey seal (Halichoerus grypus)"Kastelein, R.A., Staal, C., Wiepkema, P.R.Sensory Systems of Aquatic Mammals (Eds. Kastelein, R.A., Thomas, J.A. and Nachtigall, P.E.). De Spil Publishers, Woerden, The NetherlandsBehavior
1995"Detailed observations of suckling behaviour of a Grey seal"Kastelein, R.A., Groenenberg, H.J., Wiepkema, P.R.Int. Zoo Yb. 34, 186-200Behavior
1996"Drinking speed of Pacific walrus (Odobenus rosmarus divergens) pups"Kastelein, R.A., Postma, J., Rossum, T., Wiepkema, P.R.Aquatic Mammals 22(1), 21-26Behavior
1997"Swimming behaviour of harbour porpoises (Phocoena phocoena) under different conditions in human care"Kastelein, R.A., Staal, C.The biology of the harbour porpoise (Eds. Read, A.J., Wiepkema, P.R. and Nachtigall, P.E.). De Spil Publishers, Woerden, The Netherlands, 235-253Behavior
2004"Behaviour and Physiological effects of transmitter attachments on a captive harbour porpoise (Phocoena phocoena)"Geertsen, B.M., Teilmann, J., Kastelein, R.A., Vlemmix, H.N.J., Miller, L.A.J. Cetacean Research Management 6(2), 139-146Behavior
2015"Behavior and body mass changes of a mother and calf Pacific walrus (Odobenus rosmarus divergens) during the suckling period"Kastelein, R.A., van den Belt, I., Jennings, N., de Kruijf, R.Zoo Biology 34, 9–19Behavior
2022"White-beaked dolphins (Lagenorhynchus albirostris) cooperating with humans and showing altruism toward harbor porpoises (Phocoena phocoena)"Kastelein, R.A., Jennings, N., Huijser, L.A.E.Aquatic Mammals 48(1), 75-82, DOI 10.1578/AM.48.1.2022.75Behavior
2022"Behavioral, pathomorphological, and clinical observations of a young harbor porpoise (Phocoena phocoena) with congenital hemihydranencephaly"Kastelein, R.A., Kik, M. J. L, Huijser, L. A. E.Aquatic Mammals, 48(4), 314-323, DOI 10.1578/AM.48.4.2022.314Behavior / Pathology
1994"Oral suction of a Pacific walrus (Odobenus rosmarus divergens) in air and under water"Kastelein, R.A., Muller, M., Terlouw, A.Z. Säugetierkunde 59, 105-115Bio-mechanics
1997"Pressure changes in the mouth of a feeding harbour porpoise (Phocoena phocoena)"Kastelein, R.A., Staal, C., Terlouw, A., Muller, M.The biology of the harbour porpoise (Eds. Read, A.J., Wiepkema, P.R. and Nachtigall, P.E.). De Spil Publishers, Woerden, The Netherlands, 279-291Bio-mechanics
1990"Sensory abilities of cetaceans/laboratory and field evidence"Thomas, J.A., Kastelein, R.A.Plenum Press, New York. pp 710Co-editorship of books
1992"Sensory Systems of Marine Mammals"Thomas, J.A., Kastelein, R.A., A. SupinPlenum Press, New York. pp. 773Co-editorship of books
1995"Sensory systems of Aquatic Mammals"Kastelein, R.A., Thomas, J.A., P.E. NachtigallDe Spil Publishers, Woerden, pp. 588Co-editorship of books
1992"Cognition and decision-making by a California Sea Lion (Zalophus californianus) in visual discrimination tests"Kastelein, R.A., Mosterd, P., Vaughan, N.Sensory Systems of Marine Mammals (Eds. J.A. Thomas, R.A. Kastelein and A. Supin) Plenum Press, New York. pp. 685-700Cognition
1995"Working memory of a California sea lion (Zalophus californianus) for the 2 choice location of a 2-dimensional black stimulus"Kastelein, R.A., Herremans, A. H. J., Nieuwstraten, S. H., Wiepkema, P. R.Sensory Systems of Aquatic Mammals (Eds. Kastelein, R.A., Thomas, J.A. and Nachtigall, P.E.). De Spil Publishers, Woerden, The NetherlandsCognition
1988"Observations of Commerson's dolphin and other cetaceans in southern Chile, January-February 1984"Leatherwood, J., Kastelein, R.A., K. Miller.I.W.C. meeting Eastborne, May 1984. SC/36/SM8:71-84Ecology
1988"Studies of Commerson's dolphins, Cephalorhynchus commersonii, of Tierra del Fuego, 1976-1984, with a review of information on the species"Goodall, R.N.P., A.R. Galeazzi, Leatherwood, K.W., K.W. Miller, I.S. Cameron, Kastelein, R.A., A.P. SobralSouth Atlantic. Sc/36/SM8: 3-70Ecology
1990"Distribution, abundance, reproduction and behaviour of Steller sea lions (Eumetopias jubatus) in Prince William Sound, Alaska"Kastelein, R.A., F.C. WeltzAquatic Mammals, 15.4,145-157Ecology
1993"Information on the biology of Commerson's dolphins (Cephalorhynchus commersonii)"Kastelein, R.A., McBain, J., Neurohr, B.Aquatic Mammals 19(1), 13-19Ecology
1989"Food consumption, body measurements and weight changes of a female Killer whale (Orcinus orca)"Kastelein, R.A., Vaughan, N.Aquatic Mammals, Vol. 15.1:18-21Energetics
1990"The food consumption of Steller sea lions (Eumetopias jubatus)"Kastelein, R.A., Vaughan, N., Wiepkema, P.R.Aquatic Mammals, 15.4, 137-144Energetics
1990"The food consumption of Grey seals (Halichoerus grypus) in human care"Kastelein, R.A., Wiepkema, P.R., Vaughan, N.Aquatic Mammals 15.4,171-180Energetics
1990"The food consumption of South African fur seals (Arctocephalus pussilus) in the Harderwijk Marine Mammal Park"Kastelein, R.A., I. Verhoeven, Wiepkema, P.R.Int. Zoo Yearb. (Eds. P.J.S. Olney, P. Ellis and B. Sommerfelt) The Zool. Soc. of London. 29, 175-179Energetics
1991"The food consumption of Southern elephant seals (Mirounga leonina)"Kastelein, R.A., Kershaw, J., Wiepkema, P.R.Aquatic Mammals 17(2),76-87Energetics
1993"The food consumption of Commerson's dolphins (Cephalorhynchus commersonii)"Kastelein, R.A., McBain, J., Neurohr, B., Mohri, M., Saijo, S., Wakabayashi, I., Wiepkema, P.R.Aquat. Mamm. 19(2), 99-121Energetics
1994"Effective mass transfer in a suckling Grey seal (Halichoerus grypus)"Kastelein, R.A., Wiepkema, P.R., Berghout, E., Groenenberg, H.J.Aquatic Mammals 20(1), 11-27Energetics
1994"Food consumption, growth and reproduction of Belugas (Delphinapterus leucas) in human care"Kastelein, R.A., Ford, J., Berghout, E., Wiepkema, P.R., Boxsel, M.Aquatic Mammals 20(2), 81-97Energetics
1995"The food consumption of South American sea lions (Otaria flavescens)"Kastelein, R.A., Kershaw, J., Berghout, E., Wiepkema, P.R.Aquatic Mammals 21(1), 43-53Energetics
1997"Food consumption and body weight of harbour porpoises (Phocoena phocoena)"Kastelein, R.A., Hardeman, J., Boer, H.The biology of the harbour porpoise (Eds. Read, A.J., Wiepkema, P.R. and Nachtigall, P.E.). De Spil Publishers, Woerden, The Netherlands, 217-233Energetics
1998"Food consumption and growth of marine mammals. Ph.D. Thesis"Kastelein, R.A.De Spil Publishers, Woerden, The Netherlands, 647 ppEnergetics
1999"Food consumption and body measurements of Amazon river dolphins (Inia geoffrensis)"Kastelein, R.A., Neurohr, B., Nieuwstraten, S.H., Wiepkema, P.R.Aquatic Mammals 25.3, 173-182Energetics
2000"Food consumption, growth, body dimensions, and respiration rates of captive false killer whales (Pseudorca crassidens)"Kastelein, R.A., Mosterd, J, Schooneman, N.M., Wiepkema, P.R.Aquatic Mammals, 26.1, 33-44Energetics
2000"Food consumption and Growth of California Sea Lions (Zalophus californianus californianus)"Kastelein, R.A., Schooneman, N.M., Vaughan, N., Wiepkema, P.R.Zoo Biology 143-159Energetics
2000"Food Consumption and Growth of a female Dusky Dolphin (Lagenorhynchus obscurus)"Kastelein, R.A., Elst, van der, C.A., Tennant, H.K., Wiepkema, P.R.Zoo Biology 131-142Energetics
2000"A note on food consumption and growth of common dolphins (Delphinus delphis)"Kastelein, R.A., Macdonald, G.J., Wiepkema, P.R.J. Cetacean Research and Management 2(1),69-73Energetics
2000"Food consumption of a captive female killer whale (Orcinus orca)"Kastelein, R.A., Walton, S., Odell, D., Nieuwstraten, S.H., Wiepkema, P.R.Aquatic Mammals 26 (2), 127-131Energetics
2000"Food consumption and body weight of captive Pacific walruses (Odobenus rosmarus divergens)"Kastelein, R.A., Schooneman, N.M., Wiepkema, P.R.Aquatic Mammals 26(3), 175-190Energetics
2002"Food intake and body measurements of Atlantic bottlenose dolphins (Tursiops truncatus) in captivity"Kastelein, R.A., Vaughan, N., Walton, S., Wiepkema, P.R.Marine Environmental Research 53, 199-218Energetics
2003"Food consumption, growth and food passage times in Pacific Walrus Odobenus rosmarus divergens pups at Harderwijk Marine Mammal Park"Kastelein, R.A., Klasen, W.J.C., Postma, J., Boer, H., Wiepkema, P.R.International Zoo Yearbook, 38, 192-203Energetics
2003"Food consumption and suckling of Killer whales Orcinus orca at Marineland, Antibes"Kastelein, R.A., Kershaw, J., Berghout, E., Wiepkema, P.R.International Zoo Yearbook, 38, 204-218Energetics
2003"Food consumption, food passage time, and body measurements of captive bottlenose dolphins (Tursiops truncatus)"Kastelein, R.A., Staal, C., Wiepkema, P.R.Aquatic Mammals 29(1), 53-66Energetics
2005"Food Consumption and Body Mass of Captive Harbor Seals (Phoca vitulina)"Kastelein, R.A., Staal, C., Wiepkema, P.R.Aquatic Mammals, 31(1), 34-42Energetics
2018"Seasonal changes in food consumption, respiration rate, and body condition of a male harbor porpoise (Phocoena phocoena)"Kastelein, R.A., Helder-Hoek, L., Jennings, N.Aquatic Mammals, 44, 76-91, DOI 10.1578/AM.44.1.2018.76Energetics
2019"Reduction in body mass and blubber thickness of harbor porpoises (Phocoena phocoena) due to near-fasting for 24 hours in four seasons"Kastelein, R.A., Helder-Hoek, L., Jennings, N., van Kester, R., Huisman, R.Aquatic Mammals, 45 37-47. DOI 10.1578/AM.45.1.2019.37Energetics
2019"High levels of food intake in harbor porpoises (Phocoena phocoena): Insight into recovery from disturbance"Kastelein, R.A., Helder-Hoek, L., Booth, C., Jennings, N., Leopold, M.Aquatic Mammals, 45(4), 380-388, DOI 10.1578/AM.45.4.2019.380Energetics
1999"Inclusion of Cetaceans Within the Order Artiodactyla Based on Phylogenetic Analysis of Pancreatic Ribonuclease Genes"Kleineidam, R.G., Pesole, G., Breukelman, H.J., Beintema, J.J., Kastelein, R.A.J. Mol. Evol. 48, 360-368Genetics
1988"A safe and standardizable technique to identify individual dolphins in human care"Kastelein, R.A., T. Dokter.Aquatic Mammals, 14.1, 13-20Husbandry
1995"Improving parental care of a female bottlenose dolphin (Tursiops truncatus) by training"Kastelein, R.A., Mosterd, J.Aquatic Mammals 21, 165-169Husbandry
2001"Walrus (Odobenus rosmarus)"Kastelein, R.A.Encyclopedia of Marine Mammals. Editors: W.F. Perrin, J.G.M. Thewissen and B. Würsig. Academic press, 1294-1300Overviews
2009"Walrus (Odobenus rosmarus)"Kastelein, R.A.Encyclopedia of Marine Mammals. 2nd edition. Editors: W.F. Perrin, B. Würsig, and J.G.M. Thewissen. San Diego, Elsevier, 1212-1217Overviews
1986"Assessment of cleaning and rehabilitation procedures using live otters"Davis, R.W., Thomas, J.A., Williams, T.M., Kastelein, R.A.Pages 59-99. In Sea otter oil spill mitigation study. Report No. MMS-86-0009 to Minerals Management Service, Los AngelesPhysiology
1988"The effects of oil contamination and cleaning on sea otters (Enhydra lutris). 1. Thermoregulatory implication based on pelt studies"Williams, T.M., Kastelein, R.A., Davis, R.W., Thomas, J.A.Can. J. Zool. 66,2776-2781Physiology
1988"The effects of oil contamination and cleaning on sea otters (Enhydra lutris). II. Metabolism, thermoregulation, and behavior"Davis, R.W., Williams, T.M., Thomas, J.A., Kastelein, R.A., L.H. CornellCan. J. Zool. 66: 2782-2790. DOI: 10.13140/2.1.2317.4082Physiology
1989"Respiratory arrhytmia in the hearts of Harbour porpoises (Phocoena phocoena)"Kastelein, R.A., F.L. MeijlerAquatic Mammals, 15.4, 137-144Physiology
1990"Adaptations of marine mammals to diving"Kastelein, R.A.Proc. Kon. Ned. Akad. v. Wetensch. 93(4), 427-430Physiology
1992"Apparent digestive efficiency in walruses (Odobenus rosmarus) fed herring (Clupea harengus) and clams (Spisula sp.)"Fisher, K.I., R.E.A. Steward, Kastelein, R.A., L.D. CampbellCan. J. Zool. 70, 30-36Physiology
1997"Skin surface temperature changes in a harbour porpoise"Kastelein, R.A., Koene, P., Nieuwstraten, S.H., Labberté, S.The biology of the harbour porpoise (Eds. Read, A.J., Wiepkema, P.R. and Nachtigall, P.E.). De Spil Publishers, Woerden, The Netherlands, 255-264Physiology
1997"Passage time of carmine red dye through the digestive tract of harbour porpoises (Phocoena phocoena)"Kastelein, R.A., Nieuwstraten, S.H., Verstegen, M.W.A.The biology of the harbour porpoise (Eds. Read, A.J., Wiepkema, P.R. and Nachtigall, P.E.). De Spil Publishers, Woerden, The Netherlands, 265-275Physiology
1997"Respiration in harbour porpoises (Phocoena phocoena)"Kastelein, R.A., Vaughan, N., Groenenberg, H.J., Boekholt, H., Schreurs, V.The biology of the harbour porpoise (Eds. Read, A.J., Wiepkema, P.R. and Nachtigall, P.E.). De Spil Publishers, Woerden, The Netherlands, 203-215Physiology
2000"Gas exchange and heart rate in the harbour porpoise (Phocoena phocoena)"Reed, J. Z., Chambers, C., Hunter, C. J., Lockyer, C., Kastelein, R.A., Fedak, M. A., Boutilier, R. G.J.Comp. Physiol B 170, 1-10Physiology
2004"Behaviour and Physiological effects of transmitter attachments on a captive harbour porpoise (Phocoena phocoena)"Geertsen, B.M., Teilmann, J., Kastelein, R.A., Vlemmix, H.N.J., Miller, L.A.J. Cetacean Research Management 6(2), 139-146Physiology
1988"Estimate of numbers of Commerson's dolphins in a portion of the northeastern Strait of Magellan, January-February 1984"Leatherwood, J., Kastelein, R.A., P.S. HammondI.W.C. meeting Eastborne May 1984. SC/36/SM7:85-92Population estimates
1991"Distribution and behaviour of Steller sea lions (Eumetopias jubatus) In Prince William Sound, Alaska, June 1989"Kastelein, R.A., F.C. WeltzAquatic Mammals 17(2), 91-97Population estimates
1996"The Status of the Harbour Porpoise, Phocoena phocoena, in Dutch Waters and the State of Related Research in the Netherlands: An Overview"Reijnders, P.J.H., Leopold, M.F., Camphuysen, C.J., Heessen, H.J.L., Kastelein, R.A.Rep. Int. Whal. Commn. 46, 607-611Population estimates
1990"The medical treatment of 3 stranded Harbor porpoises (Phocoena phocoena)"Kastelein, R.A., Bakker, M., Dokter, T.Aquatic Mammals 15.4,181-202Rehabilitation of stranded cetaceans
1995"A swimming support for dolphins undergoing veterinary care"Kastelein, R.A., Dokter, T., Hilgenkamp, J.Aquatic Mammals 21(2), 155-159Rehabilitation of stranded cetaceans
1997"The rehabilitation and release of stranded harbour porpoises (Phocoena phocoena)"Kastelein, R.A., Bakker, M.J., Staal, C.The biology of the harbour porpoise (Eds. Read, A.J., Wiepkema, P.R. and Nachtigall, P.E.). De Spil Publishers, Woerden, The Netherlands, 9-61Rehabilitation of stranded cetaceans
1997"A method for tube-feeding juvenile harbour porpoises (Phocoena phocoena)"Kastelein, R.A., Schooneman, N.M., Staal, C., Boer, H.The biology of the harbour porpoise (Eds. Read, A.J., Wiepkema, P.R. and Nachtigall, P.E.). De Spil Publishers, Woerden, The Netherlands, 63-83Rehabilitation of stranded cetaceans
1999"Cutaneous Papillonmavirus infection in a harbour porpoise (Phocoena phocoena) from the North Sea"Van Bressem, M-F., Kastelein, R.A., Flamant, P., Orth, G.Veterinary Record 144, 592-593Rehabilitation of stranded cetaceans
2023"Evaluating the use of diazepam in stranded dolphins and porpoises for husbandry and veterinary purposes"Kastelein, R.A., Bakker, M.J., Jennings, N., Covi-Dijkhuizen, J.Aquatic Mammals 49(1), 94-103, DOI 10.1578/AM.49.1.2023.94Rehabilitation of stranded cetaceans
1988"The sensitivity of the vibrissae of a Pacific walrus (Odobenus rosmarus divergens). Part 1"Kastelein, R.A., van Gaalen, M.A.Aquatic Mammals, 48(1), 36-58, DOI 10.1578/AM.48.1.2022.36Sensory systems other than hearing
1990"The tactile sensitivity of the mystacial vibrissae of a Pacific Walrus (Odobenus rosmarus divergens) Part II: Masking"Kastelein, R.A., Stevens, S., Mosterd, P.Aquatic Mammals 16,78-87Sensory systems other than hearing
1990"A potential substrate for geomagnetic sensibility in cetaceans"Gerrits, N.M., Kastelein, R.A.Sensory abilities of cetaceans/Laboratory and field evidence (Eds. J.A. Thomas and R.A. Kastelein) Plenum Press, New York, 31-38Sensory systems other than hearing
1988"The significance of training for the behaviour of Steller sea lions (Eumetopias jubata) in human care"Kastelein, R.A., Wiepkema, P.R.Aquatic Mammals.14.1, 39-41Welfare (occupational therapy)
1989"The use of molluscs to occupy Pacific Walruses (Odobenus rosmarus divergens) in human care"Kastelein, R.A., Wiepkema, P.R., Slegtenhorst, C.Aquatic Mammals 15(1), 6-8Welfare (occupational therapy)
1989"The excavation technique for molluscs of Pacific Walruses (Odobenus rosmarus divergens) under controlled conditions"Kastelein, R.A., P. Mosterd, Slegtenhorst, C.Aquatic Mammals, Vol. 15.1:3-5Welfare (occupational therapy)
1990"The excavation technique for molluscs of Pacific Walruses (Odobenus rosmarus divergens) under controlled conditions"Poole, T.B., Kastelein, R.A.Ratel 17(4), 108-115Welfare (occupational therapy)
1991"Food dispensers as occupational therapy for the Walrus (Odobenus rosmarus divergens) at the Harderwijk Marine Mammal Park"Kastelein, R.A., M. Paasse, P. Klinkhamer, Wiepkema, P.R.Int. Zoo YB. 30,207-212Welfare (occupational therapy)
2007"Feeding enrichment methods for Pacific walrus (Odobenus rosmarus divergens) calves"Kastelein, R.A., Jennings, N., Postma, J.Zoo Biology 26(3), 175-186Welfare (occupational therapy)
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